NORTHERN PERU: Across the Marañon Canyon

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1 NORTHERN PERU: Across the Marañon Canyon 20 September 5 October 2015 TOUR LEADER: Nick Athanas Report by Nick Athanas; photos not otherwise credited are also Nick s. Many thanks to tour participant Eric van der Aa ( for contributing some photos used in this report Long-whiskered Owlet was a major tour highlight! Northern Peru is jam-packed with absolutely superb birds, many of which are either endemic or difficult to see elsewhere. We had great success targeting the vast majority of them on this tour, and also saw many, many other species along the way. We started in the dry Northwest with nice sightings of Peruvian Plantcutter and Rufous Flycatcher in the lowland forests and White-winged Guan and Elegant Crescentchest a bit higher up around the beautiful Chaparrí Lodge. Heading east, we stopped for Piura Chat-Tyrant and a number of dry highland specialties, and

2 then birded around Jaén for lower Marañon birds like Little Inca-Finch and Marañon Crescentchest before heading to the east slope (making an essential stop to see Marvelous Spatuletail!). With three nights each in the Waqanki Lodge at lower elevations and Owlet Lodge at higher elevations, we had time to see a great selection of humid forest birds ranging from endemics like Mishana Tyrannulet, Ash-throated Antwren, Yellow-scarfed Tanager, Chestnut Antpitta, and Long-whiskered Owlet to more widespread but memorable species like Rufous-crested Coquette, Sword-billed Hummingbird, and Paradise Tanager. A side trip to San Lorenzo was successful for Pale-billed Antpitta and other high elevation birds before we headed up the Utcubamba Valley and over the imposing east rim of the Marañon Canyon. Once again it took a while to find Yellow-faced Parrotlet, but we nailed it beautifully in the end along with the other endemics of this spectacular region. We headed to Cajamarca for the end of the tour, picking up nearly all the possible endemics in the area including Striated Earthcreeper, Gray-bellied Comet, and Great Spinetail. Weather was excellent in fact, it was arguably too good for several days on the east slope of the Andes, where hot and sunny conditions led to rather slow and quiet bird activity and few mixed species flocks, but with the prospect of a bad El Niño looming in the next couple of months, we really couldn t complain too much about having very little rain. A multinational group of friendly and excellent birders, not to mention their great senses of humor, led to a nice vibe throughout this fun an successful trip. After a single night in Lima, we took an early flight north to the desert city of Chiclayo, and immediately headed north of the city to Bosque de Pomac, a dry forest dominated by mesquite and acacia, and the largest protected forest in the region. We had to stop and register on arrival, quickly finding a number of common species like Pale-legged (Pacific) Hornero, Fasciated Wren, and Collared Antshrike, but soon headed deeper into the forest. This would be our only chance to see several key endemic birds, chief among them the endangered Peruvian Plantcutter. Last year this bird was a challenge to find, but luckily this year we did not have to walk far before hearing its odd kazoo-like calls and soon had super views of a perched male (photo left courtesy of Eric van der Aa). We also had our first sightings of other dry country species like Amazilia Hummingbird, Scarlet-backed Woodpecker, Necklaced Spinetail, Gray-and-white Tyrannulet, Superciliated Wren, and the neat-looking Tumbes Tyrant. We then drove deeper into the reserve to an area of sparser forest and sand dunes. A pair of wide-eyed Peruvian Thick-knees were trying to roost where we parked the van, and they warily trotted away as we got out, but stayed around so we could admire then in the scope. A ghostly Coastal Miner was foraging in the sand nearby, and several Tumbes Swallows circled overhead, coming lower to mob a Peruvian Pygmy-Owl that was tooting from a nearby tree. Other birds mobbed the owl including a female Short-tailed Woodstar and a few Cinereous Conebills. We were down to one major target for the area, the very localized and highly endangered Rufous Flycatcher. We chased after a distant call and eventually tracked one down for a quick view, but after that they seemed to be all over, and we saw them much better on several occasions on the walk back to the van.

3 Peruvian Thick-knee and Rufous Flycatcher at Bosque de Pomac (photos courtesy of Eric van der Aa) After a picnic lunch, we drove a couple of hours inland and up into the hills to the Chaparrí reserve, and met up with our local guide Antero. We first stopped at the Tinajones Resevoir to see Black-faced and Puna Ibises along with numerous common herons and other waterbirds, then started driving up into the reserve. We made numerous stops to see Cinereous Finch, Pacific Parrotlet, Parrot-billed Seedeater, Aplomado Falcon, and a male Short-tailed Woodstar before finally arriving at the lodge shortly before dark. White-winged Guans (photo below) greeted us as we had breakfast just before light. They are the result of an amazingly successful reintroduction project that began with 16 birds in October 2001; the population has increased to at least 100 and they are also mixing with wild birds in an adjacent canyon.

4 Feeders at the lodge attracted several birds like the gaudy White-tailed Jay along with Croaking Ground-Dove, Longtailed Mockingbird, and Groove-billed Ani, and Antero took us to a nearby territory of Elegant Crescentchest which performed well; a Gray-and-gold Warbler in the same spot was a surprise. We spent some time watching hummers come in to bathe in a stream, including Purple-collared Woodstar, Amazilia Hummingbird, and Tumbes Hummingbird, then headed out to bird some trails and tracks around the reserve. After a few hours we had added White-edged Oriole, Golden-olive Woodpecker, Red-masked Parakeet, Bran-colored Flycatcher, Tropical (Tumbes) Pewee, Short-tailed Field Tyrant, Baird s Flycatcher, Tropical Parula, Collared Warbling-Finch, Tumbes Sparrow, and Golden Grosbeak. We had a long siesta during the very hot middle of the day, and then headed back down to the flatter areas along the entrance road to search for Sulphur-throated Finch, a very localized species which is hard to find elsewhere. Finally we found a flock of mostly immature birds, though a few yellow-throated adults were also present. During our search we also saw Harris s Hawk and had our best views of Peruvian Meadowlark. At dusk, a pair of Scrub Nightjars responded nicely giving good flyover views, before we headed back to the lodge. Purple-collared Woodstar bathing in the stream at Chaparrí. Next morning we spent more time watching the hummers, where a Long-billed Starthroat was also catching insects over the water, then headed back down to Tinajones Resevoir. Stopping along the way we found a flock of seedeaters that included both Chestnut-throated and Variable Seedeaters, then tried a stakeout for Spotted Rail, which actually showed pretty well for a change, though the Many-colored Rush-Tyrant we found nearby proved to be an even bigger hit. There was a large flock of Comb Ducks at the lake, which were pretty close and in nice light. Other birds around included Cinnamon Teal, White-cheeked Pintail, Great Grebe, Black-necked Stilt, Slate-colored Coot, Hudsonian Godwit, Baird s Sandpiper, Gray-hooded Gull, and Burrowing Owl. We then headed back to Chiclayo and drove to coast, having a packed lunch on the recently renovated beachfront at Etén. Not only was it a nice spot for lunch, it proved to be great for seawatching as we scoped out flocks of Peruvian Boobies and Guanay Cormorants out near the

5 fishing boats along with Peruvian Pelicans and Kelp Gulls nearer the beach. A lone Brown Pelican was also scoped, looking small next to a Peruvian Pelican sitting next to it. We spent the afternoon birding other wetlands and beaches nearby, where shorebirds featured strongly, such as Lesser Yellowlegs, Whimbrel, Ruddy Turnstone, Sanderling, Least Sandpiper, Wilson s Phalarope, and Killdeer. We managed to tease out a Wren-like Rushbird from a large stand of reeds, and some careful searching got us a Yellowish Pipit the subspecies found here is a good candidate for a split. We spent the night at a pleasant lodge located near the pre-columbian Túcume ruins, seeing some of the old, eroded pyramids and we drove in. It was our earliest start of the trip as we hit the road at 4:15am for the long drive north then west up into the Andes to Abra Porculla, where we birded a dirt road fringed by patches of montane scrub and forest. Our excellent driver Gilmer set up the table for our field breakfast while we notched a few first birds of the morning including Plumbeous-backed Thrush and the superb Black-cowled Saltator. Breakfast always tastes better after a few nice sightings and amid the dramatic mountain scenery. We birded there for the rest of the morning, and after a slow start, activity picked up nicely, and we located a number of species that would not be possible anywhere else. Most important among them was the endemic Piura Chat-Tyrant, but we also saw Chapman s Antshrike, Line-cheeked Spinetail, Rufous-necked Foliagegleaner, Ecuadorian Piculet, White-winged and Bay-crowned Brushfinches, and Three-banded Warbler along with other more common species. After a field lunch, we continued our journey east towards Jaén, though took a detour in the afternoon to search some nice-looking desert for Little Inca-Finch. The finches decided to be shy this afternoon, and only a few of the group had a brief sighting as one perched for a half a second on a cactus, though we did get our first Spot-throated Hummingbirds and Rufous-fronted Thornbirds, and some glimpses of a Marañon Crescentchest, before driving to Jaén for the night. Ecuadorian Piculet at Abra Porculla

6 We had a lot to cram in today! Once again we were out early, driving north of Jaén to a patch of forest near the town of Tamborapa. Birds came thick and fast even before our field breakfast, and we quickly found Necklaced Spinetail (sometimes split as Chinchipe Spinetail), Northern Slaty-Antshrike, Speckle-breasted Wren, and point-blank views of Marañon Crescenchest, a far more satisfying sighting than what we had yesterday. After scarfing down breakfast, we headed up a trail and quickly connected with Black-capped Sparrow, Red-crested Finch, and Buff-bellied Tanager. Marañon Spinetails were singing but proved a nightmare to see, though some of the group managed some brief views. On a tight schedule, we left Jaén and decided to stop once again to try for Little Inca-Finch, this time finding a much more cooperative bird that perched up long enough for everyone to enjoy it. Most of the rest of the day was spent driving to Moyobamba, but we stopped to have lunch at the now-famous hummingbird feeders at Huembo, where we were blown away by the two male Marvelous Spatuletails that were regularly visiting the feeders. Other hummers there included Brown, Green and Sparkling Violetears, Bronzy Inca, Chestnut-breasted Coronet, White-bellied Hummingbird, Andean Emerald, White-bellied Woodstar, and Speckled Hummingbird. We arrived at Waqanki Lodge around sunset, having time to settle in and relax a bit after the long day. A Marvelous Spatuletail coming to a feeder at Huembo The Moyobamba area is remarkably diverse, having a mix of Andean foothill species, Amazonian species, and even wetland and savanna species, some of which have amazingly disjunct distributions. Waqanki s excellent bird guide Carlos Altamirano accompanied us during our time here, and his local knowledge helped us find a number of tough species. On our first morning we concentrated on the lower parts of the trail that goes up Quebrada Mishquiyaca, where some highlights included a gorgeous Golden-collared Toucanet, a flock of Chestnut-eared Aracaris, the beautiful little Blackand-white Tody-Flycatcher, and a cracking Fiery-capped Manakin. Antbirds also featured as we were able to call in Peruvian Warbling-Antbird, Blackish Antbird, and Spot-winged Antbird. A flock of Spot-winged Parrotlets flew over

7 very high, so it hard to appreciate this local species. Other birds seen on the walk included Blue Ground-Dove, Forktailed Palm-Swift, Yellow-tufted and Little Woodpeckers, Buff-throated Foliage-gleaner, Sooty-headed Tyrannulet, Yellow-browed Tody-Flycatcher, Piratic Flycatcher, Wing-barred Piprites, Yellow-bellied, Turquoise, Bay-headed, and Black-bellied Tanagers, Black-faced Dacnis, Purple and Green Honeycreepers, Chestnut-bellied Seed-Finch, and Russetbacked Oropendola. We spent late morning around the hummingbird feeders near the lodge, where dozens of these beautiful creatures were swarming around. Rufous-crested Coquette was a clear favorite and even perched in some nice spots for photos. Other hummers there were White-necked Jacobin, Great-billed Hermit, Black-throated Mango, Gray-breasted Sabrewing, Fork-tailed Woodnymph, Sapphire-spangled Emerald, Golden-tailed Sapphire, and Whitechinned Sapphire. In the afternoon we first drove out to a well-known stakeout for Oilbird, where a small colony roosts under a bridge near the main highway, then headed out into the savanna areas and rice paddies west of Moyobamba. A small pond had some Masked Ducks, and we tried to see a Russet-crowned Crake, though it never came into view. Carlos then took us to his stakeout for Cinereous-breasted Spinetail and Rusty-backed Antwren, both of which showed exceptionally well, even perching together in the same bush. Another stop near some roadside rice fields got us superb views of the very local Pale-eyed Blackbird as well as scope views of a distance Black-billed Seed-Finch. We spent the rest of the afternoon on some side roads through the rice, adding numerous other birds like Snail Kite, Limpkin, Wattled Jacana, Yellow-headed Caracara, Pale-legged Hornero, Black-capped Donacobius, Red-capped Cardinal, Oriole Blackbird, and Purple-throated Euphonia. Owls did not cooperate tonight at all, somewhat surprising for this area which is usually pretty good for them. Rufous-crested Coquette at the Waqanki feeders and an Oilbird under the bridge (Oilbird photo by Eric van de Aa) Next day, some of the group were keen to do the tough hike up to the ridge above Waqanki for Ash-throated Antwren, so we split up for the morning after seeing a few birds during breakfast like Crested Oropendola, Plain-brown Woodcreeper, and Black-bellied Tanager. With the threat of rain looming, Team Carlos set off straight after breakfast to get a head start up the trail, while Team Nick birded near the lodge for a while to see what the weather had in store. Luck was with us as the clouds dispersed and it ended up being a spectacular morning. Both teams had great birds

8 though quite different lists. My group first birded some scrub below the lodge and had great success in calling a Tataupa Tinamou into a small clearing, while also seeing Barred Antshrike, Short-crested Flycatcher, Dull-colored Grassquit, and Grayish Saltator nearby. Since the weather improved dramatically, we set off up the trail to look for species we hadn t seen yesterday. Activity was nice and we had great birds like Bluish-fronted Jacamar, Gilded Barbet, Blue-headed Parrot perched very close, very cooperative Stripe-chested Antwren and Spot-backed Antbird, an early Cerulean Warbler, the endemic Mishana Tyrannulet, Gray-headed Kite, Short-tailed Hawk, Olivaceous Woodcreeper, Whitelored Tyrannulet, Euler s Flycatcher, Coraya Wren, Yellow-crested Tanager, Paradise Tanager, and White-vented Euphonia. One clearing was full of trogons calling back and forth and we had males and females of both Green-backed and Blue-crowned Trogons. We had just finished our lunch back at the lodge when a tired but elated Team Carlos arrived back with news of point-blank views of Ash-throated Antwren! Despite their much longer walk, they also managed to see a nice selection of other birds like Gilded Barbet, Plain-winged Antshrike, Plain Antvireo, Ocellated Woodcreeper, Chestnut-throated Spinetail, Red-billed and Mishana Tyrannulets, White-throated Spadebill, Olivechested Flycatcher, Fiery-throated Fruiteater, White-winged Becard, Varzea Thrush, Paradise Tanager, Swallow Tanager, and Blue-naped Chlorophonia. In the afternoon, we spent an hour or so birding the base of a nearby mountain called Morro de Calzada, where we managed to see Pearly-vented and Stripe-necked Tody-Tyrants, Plumbeous Kite, Forest Elaenia, some migrating Eastern Kingbirds, and a Fork-tailed Flycatcher, which is pretty unusual in this location. We then spent the rest of the afternoon at the hummingbird feeders, where Black-throated Hermit, Amethyst Woodstar, and Violet-fronted Brilliant finally put in appearances. Our poor owling luck continued that evening when a stakeout for Black-banded failed to produce and Band-bellied stayed silent, though we did enjoy watching the impressive Supermoon rise over the forest and got to see some of the total lunar eclipse. Violet-fronted Brilliant Black-throated Hermit

9 Moonrise over Waqanki Lodge We woke up to the otherworldly calls of a Band-bellied Owl resonating through the windows, and this time we finally connected as the bird actually came in and perched in a nearby tree. Even more amazingly, the whole group had managed to get dressed and grab bins fast enough to see it! After starting the trip the day with an owl, we were sincerely hoping to end it with one too, but there was a lot to do first. After breakfast we birded around the lodge a bit, finding another Bluish-fronted Jacamar, Wedge-billed Woodcreeper, and Buff-rumped Warbler before heading back west. Hilmar, our driver, did a great job spotting Spot-breasted Woodpecker and Lettered Aracari along the side of the highway before we arrived at the Arena Blanca reserve, whose hummer feeders were worth a visit for both Green and Blue-fronted Lancebills, Wire-crested Thorntail (photo below), Many-spotted Hummingbird, and Long-billed Starthroat.

10 Other birds around Arena Blanca included Black-faced Tanager, Dark-breasted Spinetail, Olive-striped Flycatcher, and Gray-capped Flycatcher. As we worked out way up to Abra Patricia and Owlet Lodge, we made various stops along the road, hoping to find some of the superb mixed species flocks the area is known for. Hot and sunny weather made the birding fairly quiet, and we only had a small flocks with Slaty-capped Flycatcher, Three-striped Warbler, Ash-browed Spinetail, and Streaked Xenops. However we were successful finding a beautiful male Royal Sunangel, and in the same spot a flock of noisy White-capped Tanagers came in very close and screamed at us. Arriving at Owlet Lodge, we spent time at the feeders seeing new hummingbirds like Emerald-bellied Puffleg, Long-tailed Sylph, Fawn-breasted Brilliant, and Collared Inca, and also saw Peruvian Tyrannulet, Grass-green Tanager, and Bluish Flowerpiercer near the lodge. Long-tailed Sylph at the Owlet Lodge feeders (photo courtesy of Eric van der Aa). The weather was starting to look at little ominous, but we decided to still make an attempt to see the Long-whiskered Owlet. For all we knew, we could get rained out the other evenings, so we had to try. It s about a 1.2 km walk down into a valley to the most reliable spot, and Roberto, one of the reserve s rangers, knew exactly where to look. We took our time on the way down and had some sightings like Yellow-scarfed Tanager (though we would see it better later), Inca Flycatcher, Black-capped Hemispingus, and Striped Treehunter. A Rufous-bellied Nighthawk flew over at dusk giving loud whistles, and then it was time. Roberto showed us where to stand and he started calling the owlet. It started to sprinkle... were we going to get rained out now? Luckily it was only a few drops and after a few minutes we heard the soft calls of the Long-whiskered Owlet answering. It sounded a bit closer, but Robert was patient. Finally it sounded as if it were right overhead, and Roberto turned on the spotlight there it was! Very close views of this unique owl, which was only described in 1977 and virtually unknown in life until the lodge was built in The great sighting made the long walk back up the slope much easier to bear the lodge staff even made us some Pisco Sours to toast with. The little rain we had the previous night was a tease, and most of the next two days were clear, hot, and sunny, which is pretty unusual for this normally wet region. Unfortunately, the birds here are happier when it is wet, so the going was a bit tough. There were times when we walked down trails in prime morning hours and heard nothing but insects.

11 However with patience and persistence, we did manage to see most of the regional specialties along with a good selection of east slope Andean cloudforest species. On our first morning, we started out birding the entrance road and the highway near the lodge, successfully finding the endemic Johnson s Tody- Flycatcher (photo right) along with White-throated Quail-Dove, Pale-edged Flycatcher, Green-and-black Fruiteater, Sharpe s and Mountain Wrens, Spectacled Whitestart, Blue-capped Tanager, and Yellow-breasted Brush-Finch before driving down the road to a different trail. We got held up at some road works where the work crew were, no joke, waiting for paint to dry! We did get to see Tyrannine Woodcreeper and Tawnybellied Hermit during the delay. Once we got to the trail it was hot and quiet, and almost none of our targets performed, though some of the group did get some views of Rufous-vented Tapaculo. Another nearby trail was a little better giving us some decent looks at Bar-winged Wood-Wren. In the afternoon, we worked trails behind the lodge, which again were really slow for a while (though we did find a Variable Antshrike) until a nice mixed-species flock really livened things up and the trees overhead filled up with birds like Pearled Treerunner, Sulphur-bellied and Mottle-cheeked Tyrannulets, Cinnamon Flycatcher, Blue-and-black, Beryl-spangled, Saffron-crowned, and Flame-faced Tanagers, Capped Conebill, and Chestnut-breasted Chlorophonia. A Yungas Pygmy-Owl was singing nearby, but we only saw it fly away. Taking a different trail back to the lodge, we spent time trying to see Rusty-tinged Antpitta, which circled around and darted across the trail a couple of times. Back at the lodge for dusk, we enjoyed seeing a Sword-billed Hummingbird visit the feeder as well as a couple of mammals, a very cool-looking Tayra and a Black Agouti. A giant flock of Scaly-naped Parrots flew low overhead as well. We hit the trails again early next morning, this time having superb views of the endemic Chestnut Antpitta, which paused in the trail long enough for everyone to see. Once again we headed down the road (unhindered this time, at least for the moment...), birding a different reserve which was shockingly absolutely dead despite the early hour. Cinnamon-breasted Tody-Tyrant called but refused to come in, though the hummer feeders did get us some male Booted Racket-tails and a Greenish Puffleg (note that most references claim that Buff-thighed Puffleg is the species occurring in northern Peru, however recent evidence, not to mention the pure white puffs evident in the bird we saw, see photo left, indicate that it is indeed Greenish Puffleg, not Buff-thighed). We did also manage to see a male Uniform Antshrike near the feeders. Hoping for better activity down lower, we drove down into the foothills, though once again got held

12 up by the road painting crew, this time seeing Cliff Flycatcher during the delay. Clouds were starting to build as we birded the roadsides near Afluente, and we had some decent mixed-species flocks with Versicolored Barbet, Lined Antshrike, Yellow-breasted Antwren, Ecuadorian Tyrannulet, Olivaceous Siskin, and a slew of tanagers: Orange-eared, Blue-necked, Spotted, Golden-eared, Green-and-gold, and Golden. A little gap on the side of the road was enough to call in a White-crowned Tapaculo. Finally the skies opened up and rain poured down for an hour or so. When it stopped, it was cool and misty, and suddenly, just like that, birds were everywhere! We started with Rufous-tailed Tyrant at a covered scenic viewpoint as the rain was letting up, then birded a different trail for Emerald Toucanet, Streak-headed Antbird, Olive-backed Woodcreeper, Streak-necked Flycatcher, Green Jay, Russet-crowned Warbler, Oleaginous Hemispingous, Blue-winged Mountain-Tanager, Yellow-throated Tanager, Deep-blue Flowerpiercer, Golden-naped Tanager, and Silvery Tanager. Back up near the lodge activity was still good with Montane Woodcreeper, Buff-browed Foliage-gleaner, and Barred Becard. An optional late evening owling excursion was a great success with superb closeups of Cinnamon Screech-Owl. Cinnamon Screech-Owl from the trails at Owlet Lodge

13 We packed up and left Owlet Lodge early next morning, driving about an hour to the village of San Lorenzo. We hiked up a steep trail through small patches of temperate forest, frequently stopping to catch our breath in the rarified air. Tyrian Metaltail, Violet-throated Starfrontlet, Mountain Caracara, Speckle-faced Parrot, and Rufous-capped Antshrike all showed themselves during our ascent before we finally reached the bamboo-laden forest where we might find Pale-billed Antpitta. Scrambling down off the trail, we found a nice clearing and started calling. Soon the antpitta bounced past, giving our only views of this tough endemic. Back up on the trail, the endemic Russet-mantled Softtail responded almost immediately coming in close. We tried another side track which got us into the bamboo, and managed to see a pair of the local subspecies of Plain-tailed Wren, a likely split with a truly fantastic song. Both Blackish and Trilling Tapaculos were singing, but only Trilling showed itself, and we had a few Citrine Warblers. On the way back down we spotted a soaring White-throated Hawk along with Redcrested Cotinga, Rufous-breasted Chat-Tyrant, and White-sided Flowerpiercer. That afternoon, we drove south through the scenic Utcubamba Canyon, seeing Little Woodstar, Black-necked Woodpecker, Black Phoebe, Specklebreasted Wren, Marañon Thrush, and Silvery Tanager. Speckle-chested Piculet (photo upper left) took some time, but finally we had point-blank views of three of this superb and very localized endemic. We arrived in Leymebamba with barely enough time to get Rainbow Starfrontlet at some hummer feeders, where we also saw a Rufous-chested Tanager. We had one night in a very nice and creatively-decorated hotel in the town. First light saw us at some forest patches above Leymebamba, and we were greeted by a chattering Andean Flicker while Brown-bellied Swallows circled overhead. Flowerpiercers and hummers were all over and we quickly found Green-tailed Trainbearer, Shining Sunbeam, Mountain Velvetbreast, and Masked, Moustached, and Black-throated Flowerpiercers. We then lucked into a superb mixed species flock with beauties like Scarlet-bellied Mountain-Tanager, Yellow-scarfed Tanager (photo right), and Blue-backed Conebill along with more somberly-colored birds such as White-banded and Whitethroated Tyrannulets, Sierran Elaenia, and Drab Hemispingus. Heading up the road, we stopped for Andean Lapwings strutting around a cow pasture, a soaring Black-chested Buzzard-Eagle, a pair of Brown-backed Chat-Tyrants, and a Plain-colored Seedeater, before reaching some gnarled, treeline forest. Weather was surprisingly pleasant (it usually isn t..), and we soon found the endemic Coppery Metaltail

14 along with Sapphire-vented Puffleg, a fierce-looking Yungas Pygmy-Owl, Black-capped Tyrannulet, and White-chinned Thistletail. Finally we reached the 3600 m. (11800 ft.) pass (Abra Barro Negro, or Black Mud Pass) at the rim of the impressive Marañon Canyon, which is far deeper than the Grand Canyon of the US. Remarkably nice weather allowed us to enjoy the view and a few birds like Great Sapphirewing, Tufted Tit-Tyrant, and Plumbeous Sierra-Finch. After a picnic lunch, we began an incredible descent down the now-paved road towards the bottom of the canyon, stopping occasionally for scenery photos. Layers came off fast at the temperature skyrocketed, and we were all down to t-shirts by the time we stopped to see one of our targets, the fantastic Buff-bridled Inca-Finch. Some Bare-faced Ground-Doves burst off the road and disappeared before anyone really saw them this would turn out to be very fortunate... As we reached the river, it was blistering outside, and after making sure we saw some Peruvian Pigeons, we enjoyed cold drinks in the town of Balsas, then crossed the river and drove up into the shade of the western side of the canyon. In the late afternoon we had about an hour or so to look for Yellow-faced Parrotlet, the most difficult of the Marañon Canyon endemics. We heard it couple of times distantly, but it never showed itself, so we decided to leave it until the next morning and drove up over the west rim of the canyon. A White-winged Black-Tyrant was perched near the side of the road part way up, but we didn t see much else before arriving in the bustling town of Celendín, where we d spend the next two nights. Coppery Metaltail near Abra Barro Negro. Photo courtesy of Eric van der Aa. Dawn found us back down in the desert where we had heard the parrotlets the previous afternoon, but once again they were calling very far away and we could not pick them out, though we did have the tour s only Yellow-tailed Oriole. It was getting later in the morning, and I was worried the other endemics we still needed to look for would become harder to find, so we slowly started driving back up the road. A few Bare-faced Ground-Doves flew up of the road and landed in a tree, and since we had only glimpsed them the day before, we all got out. As we were looking at the doves, several Yellow-faced Parrotlets started calling nearby, and very soon we located them, for unmatched views of this beautiful little parrot. Thank you ground-doves! Spirits were high after that, and we had no difficulty seeing the other two endemics of this area: Chestnut-backed Thornbill and Gray-winged Inca-Finch. A Masked Yellowthroat was also a nice prize; the subspecies here (peruviana) is candidate for a split. Closer to Celendín, we stopped to see Jelski s Chat-Tyrant,

15 Baron s Spinetail, and the baroni subspecies of Yellow-breasted Brush-Finch. We had a pretty easy afternoon, mostly catching up on rest, and the little birding we did do did not produce any new trip birds. Three of the Yellow-faced Parrotlets we saw in the Marañon Canyon The road between Celendín and Cajamarca was paved a few years back, but it s still great birding, and most of the birds are very easy to see. Even before our field breakfast we had already managed to see one of our top targets, the endemic Striated Earthcreeper, along with some Black-crested Warblers and the first of several Yellow-billed Tit-Tyrants. Breakfast was interrupted by some Peruvian Sierra-Finches, but we would see many more through the morning. After the field breakfast, we walked along the highway finding Andean Gull, Rusty-crowed Tit-Spinetail, Black-billed Shrike- Tyrant, Tit-like Dacnis, and Golden-billed Saltator (photo left). Looking down a steep slope we managed to spot Black-crested Tit-Tyrant and White-browed Chat-Tyrant at a great distance, but we would see both much better by the end of the day. Not having found a White-tailed Shrike-Tyrant yet, I thought we should backtrack a little to an old stakeout for it, but we only had to go a couple of miles before one was perched on wire next to a Rufous-webbed Bush-Tyrant; as we watched them, a huge flock of Puna Ibis circled around and flew back in forth in a chaotic pattern. Continuing on towards Cajamarca, we made a few stops to see Black Metaltail, Slender-billed Miner, Cream-winged Cinclodes, and Streak-throated Canastero before skirting the edge of the city and driving up into the Rio Chonta Canyon. We managed to find the endemic Gray-bellied Comet in record time along with the dipperlike White-winged Cinclodes, Torrent Tyrannulet, Streak-throated Bush-Tyrant, and Band-tailed Seedeater before heading to our nice hotel right on the main square in the city center.

16 Black-crested Tit-Tyrant White-tailed Shrike-Tyrant White-browed Chat-Tyrant. Photo courtesy of Eric van der Aa With only one morning left on the trip, we made the most of it and once again left in the dark, heading back east to a valley blanketed with dense arid scrub and woodland. Our target for the morning was the very localized Great Spinetail, and with a bit of effort and some bushwhacking we managed to see a pair of them before our final field breakfast. With some extra time available, I decided to head up to Quellyacocha, a lake I had never had a chance to visit, and it turned out to be a productive side trip. Puna Teal was a nice surprise, near the northern limit of its range, while Yellow-billed Pintail and Yellow-billed Teal were also present. A Plumbeous Rail started singing from the reeds, and with a bit of playback it decided to come out to the edge. We also had a distant Silvery Grebe and a better view of Slender-billed Miner before heading back to Cajamarca. We had time to get packed up and have a nice long lunch before heading to the airport and flying back to Lima, where the tour ended. Some of the group had arranged an extra night in Lima for some sightseeing, while the rest of us caught late evening flights home. Many thanks to all for joining this excellent trip and I hope to bird with you again in the future!

17 BIRD LIST Tropical Birding Trip Report Northern Peru, September-October 2015 This list includes all the bird species that were recorded by at least one of the group, including the leader. Taxonomy and nomenclature follow Clements/eBird, August 2015 version. (Online version available here: Totals: 510 bird species seen 52 heard only H=Heard only GO=Guide only TINAMOUS TINAMIDAE H Cinereous Tinamou Crypturellus cinereus H Little Tinamou Crypturellus soui Tataupa Tinamou Crypturellus tataupa DUCKS, GEESE, AND WATERFOWL ANATIDAE Comb Duck Sarkidiornis melanotos Cinnamon Teal Anas cyanoptera White-cheeked Pintail Anas bahamensis Yellow-billed Pintail Anas georgica Puna Teal Anas puna Yellow-billed Teal Anas flavirostris Masked Duck Nomonyx dominicus CRACIDS CRACIDAE H Speckled Chachalaca Ortalis guttata White-winged Guan (E) Penelope albipennis GREBES PODICIPEDIDAE Great Grebe Podiceps major Silvery Grebe Podiceps occipitalis juninensis BOOBIES AND GANNETS SULIDAE Peruvian Booby Sula variegata CORMORANTS AND SHAGS PHALACROCORACIDAE Neotropic Cormorant Phalacrocorax brasilianus Guanay Cormorant Phalacrocorax bougainvillii PELICANS PELECANIDAE Brown Pelican Pelecanus occidentalis Peruvian Pelican Pelecanus thagus HERONS, EGRETS, AND BITTERNS ARDEIDAE Fasciated Tiger-Heron Tigrisoma fasciatum Cocoi Heron Ardea cocoi Great Egret Ardea alba Snowy Egret Egretta thula Little Blue Heron Egretta caerulea Cattle Egret Bubulcus ibis Striated Heron Butorides striata

18 IBISES AND SPOONBILLS THRESKIORNITHIDAE Puna Ibis Plegadis ridgwayi Black-faced Ibis Theristicus melanopis NEW WORLD VULTURES CATHARTIDAE Black Vulture Coragyps atratus Turkey Vulture Cathartes aura Greater Yellow-headed Vulture Cathartes melambrotus OSPREY PANDIONIDAE Osprey Pandion haliaetus HAWKS, EAGLES, AND KITES BURHINIDAE Gray-headed Kite Leptodon cayanensis Swallow-tailed Kite Elanoides forficatus Snail Kite Rostrhamus sociabilis Plumbeous Kite Ictinia plumbea Sharp-shinned (Plain-breasted) Hawk Accipiter striatus ventralis Savanna Hawk Buteogallus meridionalis Roadside Hawk Rupornis magnirostris Harris's Hawk Parabuteo unicinctus Variable Hawk Geranoaetus polyosoma Black-chested Buzzard-Eagle Geranoaetus melanoleucus Short-tailed Hawk Buteo brachyurus White-throated Hawk Buteo albigula RAILS, GALLINULES, AND COOTS HAEMATOPODIDAE H Russet-crowned Crake Anurolimnas viridis Spotted Rail Pardirallus maculatus Plumbeous Rail Pardirallus sanguinolentus Purple Gallinule Porphyrio martinicus Common Gallinule Gallinula galeata Slate-colored (Andean) Coot Fulica ardesiaca LIMPKIN JACANIDAE Limpkin Aramus guarauna THICK-KNEES THINOCORIDAE Peruvian Thick-knee Burhinus superciliaris STILTS AND AVOCETS Coragyps atratus Black-necked Stilt Himantopus mexicanus PLOVERS AND LAPWINGS LARIDAE Black-bellied (Gray) Plover Pluvialis squatarola Andean Lapwing Vanellus resplendens Killdeer Charadrius vociferus JACANAS Cathartes aura Wattled Jacana Jacana jacana SANDPIPERS AND ALLIES Cathartes burrovianus Spotted Sandpiper Actitis macularius Solitary Sandpiper Tringa solitaria Lesser Yellowlegs Tringa flavipes Whimbrel Numenius phaeopus Hudsonian Godwit Limosa haemastica Ruddy Turnstone Arenaria interpres Sanderling Calidris alba Baird's Sandpiper Calidris bairdii Least Sandpiper Calidris minutilla

19 Wilson's Phalarope Phalaropus tricolor GULLS, TERNS, AND SKIMMERS Vultur gryphus Andean Gull Chroicocephalus serranus Gray-hooded Gull Chroicocephalus cirrocephalus Kelp Gull Larus dominicanus PIGEONS AND DOVES COLUMBIDAE Rock Pigeon Columba livia Band-tailed Pigeon Patagioenas fasciata Peruvian Pigeon (E) Patagioenas oenops H Plumbeous Pigeon Patagioenas plumbea Ruddy Pigeon Patagioenas subvinacea Ruddy Ground-Dove Columbina talpacoti Croaking Ground-Dove Columbina cruziana Blue Ground-Dove Claravis pretiosa Bare-faced Ground-Dove Metriopelia ceciliae White-tipped Dove Leptotila verreauxi Gray-fronted Dove Leptotila rufaxilla White-throated Quail-Dove Zentrygon frenata West Peruvian Dove Zenaida meloda Eared Dove Zenaida auriculata CUCKOOS CUCULIDAE Squirrel Cuckoo Piaya cayana Striped Cuckoo Tapera naevia Smooth-billed Ani Crotophaga ani Groove-billed Ani Crotophaga sulcirostris BARN-OWLS TYTONIDAE Barn Owl Tyto alba OWLS STRIGIDAE H Tropical Screech-Owl Megascops choliba H Peruvian Screech-Owl Megascops roboratus Cinnamon Screech-Owl Megascops petersoni GO White-throated Screech-Owl Megascops albogularis Band-bellied Owl Pulsatrix melanota Yungas Pygmy-Owl Glaucidium bolivianum Peruvian Pygmy-Owl Glaucidium peruanum Long-whiskered Owlet (E) Xenoglaux loweryi Burrowing Owl Athene cunicularia H Rufous-banded Owl Ciccaba albitarsis NIGHTJARS AND ALLIES CAPRIMULGIDAE Rufous-bellied Nighthawk Lurocalis rufiventris Common Pauraque Nyctidromus albicollis Scrub Nightjar Nyctidromus anthonyi H Rufous Nightjar Antrostomus rufus OILBIRD STEATORNITHIDAE Oilbird Steatornis caripensis SWIFTS APODIDAE White-collared Swift Streptoprocne zonaris White-tipped Swift Aeronautes montivagus Fork-tailed Palm-Swift Tachornis squamata HUMMINGBIRDS TROCHILIDAE White-necked Jacobin Florisuga mellivora

20 Green Hermit Phaethornis guy Tawny-bellied Hermit Phaethornis syrmatophorus Great-billed Hermit Phaethornis malaris Black-throated Hermit Phaethornis atrimentalis Green-fronted Lancebill Doryfera ludovicae Blue-fronted Lancebill Doryfera johannae Brown Violetear Colibri delphinae Green Violetear Colibri thalassinus Sparkling Violetear Colibri coruscans Black-throated Mango Anthracothorax nigricollis Amethyst-throated Sunangel Heliangelus amethysticollis Royal Sunangel Heliangelus regalis Wire-crested Thorntail Discosura popelairii Rufous-crested Coquette Lophornis delattrei Speckled Hummingbird Adelomyia melanogenys Long-tailed Sylph Aglaiocercus kingii Gray-bellied Comet (E) Taphrolesbia griseiventris Green-tailed Trainbearer Lesbia nuna Tyrian Metaltail Metallura tyrianthina Coppery Metaltail (E) Metallura theresiae Black Metaltail (E) Metallura phoebe Greenish Puffleg Haplophaedia aureliae Sapphire-vented (Coppery-naped) Puffleg Eriocnemis luciani catharina Emerald-bellied Puffleg Eriocnemis aline Marvelous Spatuletail (E) Loddigesia mirabilis Shining Sunbeam Aglaeactis cupripennis Bronzy Inca Coeligena coeligena Collared Inca Coeligena torquata Violet-throated Starfrontlet Coeligena violifer Rainbow Starfrontlet Coeligena iris Mountain Velvetbreast Lafresnaya lafresnayi Sword-billed Hummingbird Ensifera ensifera Great Sapphirewing Pterophanes cyanopterus Chestnut-breasted Coronet Boissonneaua matthewsii Booted Racket-tail Ocreatus underwoodii Fawn-breasted Brilliant Heliodoxa rubinoides Violet-fronted Brilliant Heliodoxa leadbeateri Long-billed Starthroat Heliomaster longirostris Purple-collared Woodstar Myrtis fanny White-bellied Woodstar Chaetocercus mulsant Little Woodstar Chaetocercus bombus Short-tailed Woodstar Myrmia micrura Amethyst Woodstar Calliphlox amethystina Blue-tailed Emerald Chlorostilbon mellisugus Gray-breasted Sabrewing Campylopterus largipennis Fork-tailed Woodnymph Thalurania furcata Many-spotted Hummingbird Taphrospilus hypostictus Tumbes Hummingbird Leucippus baeri Spot-throated Hummingbird Leucippus taczanowskii White-bellied Hummingbird Amazilia chionogaster Amazilia Hummingbird Amazilia amazilia

21 Andean Emerald Amazilia franciae Sapphire-spangled Emerald Amazilia lactea Golden-tailed Sapphire Chrysuronia oenone White-chinned Sapphire Hylocharis cyanus TROGONS TROGONIDAE H Golden-headed Quetzal Pharomachrus auriceps Green-backed (Amaz. White-tailed) Trogon Trogon viridis Blue-crowned Trogon Trogon curucui H Collared Trogon Trogon collaris H Masked Trogon Trogon personatus MOTMOTS MOMOTIDAE H Rufous Motmot Baryphthengus martii KINGFISHERS ALCEDINIDAE Ringed Kingfisher Megaceryle torquata GO Green Kingfisher Chloroceryle americana JACAMARS GALBULIDAE Bluish-fronted Jacamar Galbula cyanescens NEW WORLD BARBETS CAPITONIDAE Gilded Barbet Capito auratus Versicolored Barbet Eubucco versicolor TOUCANS RAMPHASTIDAE Emerald (Black-throated) Toucanet Aulacorhynchus prasinus Lettered Aracari Pteroglossus inscriptus Chestnut-eared Aracari Pteroglossus castanotis Golden-collared Toucanet Selenidera reinwardtii H Yellow-throated (Black-mandibled) Toucan Ramphastos ambiguus H Channel-billed Toucan Ramphastos vitellinus WOODPECKERS PICIDAE Ecuadorian Piculet Picumnus sclateri Speckle-chested Piculet (E) Picumnus steindachneri Yellow-tufted Woodpecker Melanerpes cruentatus Little Woodpecker Veniliornis passerinus Scarlet-backed Woodpecker Veniliornis callonotus Golden-olive Woodpecker Colaptes rubiginosus H Crimson-mantled Woodpecker Colaptes rivolii Black-necked Woodpecker (E) Colaptes atricollis Spot-breasted Woodpecker Colaptes punctigula Andean Flicker Colaptes rupicola H Lineated Woodpecker Dryocopus lineatus FALCONS AND CARACARAS FALCONIDAE Black Caracara Daptrius ater Mountain Caracara Phalcoboenus megalopterus Crested Caracara Caracara cheriway Yellow-headed Caracara Milvago chimachima H Laughing Falcon Herpetotheres cachinnans American Kestrel Falco sparverius Aplomado Falcon Falco femoralis PARROTS PSITTACIDAE Spot-winged Parrotlet Touit stictopterus Cobalt-winged Parakeet Brotogeris cyanoptera Speckle-faced (White-capped) Parrot Pionus tumultuosus seniloides

22 Blue-headed Parrot Pionus menstruus Scaly-naped Parrot Amazona mercenarius Pacific Parrotlet Forpus coelestis Yellow-faced Parrotlet (E) Forpus xanthops Scarlet-fronted Parakeet Psittacara wagleri Red-masked Parakeet Psittacara erythrogenys White-eyed Parakeet Psittacara leucophthalmus TYPICAL ANTBIRDS THAMNOPHILIDAE Fasciated Antshrike Cymbilaimus lineatus Barred Antshrike Thamnophilus doliatus Rufous-capped Antshrike Thamnophilus ruficapillus Chapman's Antshrike Thamnophilus zarumae Lined Antshrike Thamnophilus tenuepunctatus Collared Antshrike (Tumbesian) Thamnophilus bernardi bernardi H Collared Antshrike (Marañon) Thamnophilus bernardi shumbae Plain-winged Antshrike Thamnophilus schistaceus Northern Slaty-Antshrike Thamnophilus punctatus leucogaster Variable Antshrike Thamnophilus caerulescens subandinus Uniform Antshrike Thamnophilus unicolor Plain Antvireo Dysithamnus mentalis H Foothill Antwren Epinecrophylla spodionota Stripe-chested Antwren Myrmotherula longicauda Gray Antwren Myrmotherula menetriesii Ash-throated Antwren (E) Herpsilochmus parkeri Yellow-breasted Antwren Herpsilochmus axillaris Rusty-backed Antwren Formicivora rufa Streak-headed (Long-tailed) Antbird Drymophila striaticeps Peruvian Warbling-Antbird Hypocnemis peruviana Blackish Antbird Cercomacroides nigrescens White-backed Fire-eye Pyriglena leuconota Spot-winged Antbird Schistocichla leucostigma Spot-backed Antbird Hylophylax naevius CRESCENTCHESTS MELANOPAREIIDAE Marañon Crescentchest Melanopareia maranonica Elegant Crescentchest Melanopareia elegans ANTPITTAS GRALLARIIDAE H Undulated Antpitta Grallaria squamigera H Scaled Antpitta Grallaria guatimalensis H Chestnut-crowned Antpitta Grallaria ruficapilla Pale-billed Antpitta (E) Grallaria carrikeri Rusty-tinged Antpitta (E) Grallaria przewalskii H Rufous Antpitta (undescribed form) Grallaria rufula (tax. nov.) H Rufous Antpitta (Cajamarca) Grallaria rufula cajamarcae Chestnut Antpitta (E) Grallaria blakei H Ochre-fronted Antpitta (E) Grallaricula ochraceifrons H Rusty-breasted Antpitta Grallaricula ferrugineipectus TAPACULOS RHINOCRYPTIDAE H Blackish Tapaculo Scytalopus latrans intermedius Trilling Tapaculo Scytalopus parvirostris Rufous-vented Tapaculo (E) Scytalopus femoralis White-crowned Tapaculo Scytalopus atratus

23 H Neblina Tapaculo (E) Scytalopus altirostris ANTTHRUSHES FORMICARIIDAE H Short-tailed Antthrush Chamaeza campanisona H Barred Antthrush Chamaeza mollissima OVENBIRDS AND WOODCREEPERS FURNARIIDAE Coastal Miner (E) Geositta peruviana Slender-billed Miner Geositta tenuirostris Olivaceous Woodcreeper Sittasomus griseicapillus amazonus Tyrannine Woodcreeper Dendrocincla tyrannina Plain-brown Woodcreeper Dendrocincla fuliginosa Wedge-billed Woodcreeper Glyphorynchus spirurus Black-banded Woodcreeper Dendrocolaptes picumnus Strong-billed Woodcreeper Xiphocolaptes promeropirhynchus Ocellated Woodcreeper Xiphorhynchus ocellatus chunchotambo Olive-backed Woodcreeper Xiphorhynchus triangularis Streak-headed Woodcreeper Lepidocolaptes souleyetii Montane Woodcreeper Lepidocolaptes lacrymiger H Duida (Lineated) Woodcreeper Lepidocolaptes duidae Plain Xenops Xenops minutus Streaked Xenops Xenops rutilans H Streaked Tuftedcheek Pseudocolaptes boissonneautii Pale-legged Hornero Furnarius leucopus tricolor Pale-legged (Pacific) Hornero Furnarius leucopus cinnamomeus Wren-like Rushbird Phleocryptes melanops Striated Earthcreeper (E) Geocerthia serrana Cream-winged Cinclodes Cinclodes albiventris White-winged Cinclodes Cinclodes atacamensis Buff-browed Foliage-gleaner Syndactyla rufosuperciliata Rufous-necked Foliage-gleaner Syndactyla ruficollis H Henna-hooded Foliage-gleaner Clibanornis erythrocephalus Striped Treehunter Thripadectes holostictus Buff-throated Foliage-gleaner Automolus ochrolaemus Pearled Treerunner Margarornis squamiger Rusty-crowned Tit-Spinetail (E) Leptasthenura pileata cajabambae Rufous-fronted Thornbird Phacellodomus rufifrons peruvianus Chestnut-backed Thornbird (E) Phacellodomus dorsalis Many-striped Canastero Asthenes flammulata Streak-throated Canastero Asthenes humilis White-chinned Thistletail Asthenes fuliginosa peruviana Russet-mantled Softtail (E) Thripophaga berlepschi Ash-browed Spinetail Cranioleuca curtata Line-cheeked Spinetail Cranioleuca antisiensis Baron's Spinetail (E) Cranioleuca baroni Azara's Spinetail Synallaxis azarae Dark-breasted Spinetail Synallaxis albigularis Cinereous-breasted Spinetail Synallaxis hypospodia Chestnut-throated Spinetail Synallaxis cherriei H Rufous Spinetail Synallaxis unirufa Marañon Spinetail Synallaxis maranonica Great Spinetail (E) Synallaxis hypochondriaca Necklaced Spinetail Synallaxis stictothorax maculata

24 Necklaced (Chinchipe) Spinetail Synallaxis stictothorax chinchipensis TYRANT FLYCATCHERS TYRANNIDAE White-lored Tyrannulet Ornithion inerme Southern Beardless-Tyrannulet Camptostoma obsoletum White-banded Tyrannulet Mecocerculus stictopterus White-throated Tyrannulet Mecocerculus leucophrys Sulphur-bellied Tyrannulet Mecocerculus minor Black-crested Tit-Tyrant Anairetes nigrocristatus Yellow-billed Tit-Tyrant Anairetes flavirostris Tufted Tit-Tyrant Anairetes parulus Mouse-colored (Tumbesian) Tyrannulet Phaeomyias murina tumbezana/inflava Mouse-colored Tyrannulet Phaeomyias murina wagae Yellow Tyrannulet Capsiempis flaveola Yellow-crowned Tyrannulet Tyrannulus elatus Forest Elaenia Myiopagis gaimardii Yellow-bellied Elaenia Elaenia flavogaster White-crested Elaenia Elaenia albiceps Sierran Elaenia Elaenia pallatangae Torrent Tyrannulet Serpophaga cinerea Streak-necked Flycatcher Mionectes striaticollis Olive-striped Flycatcher Mionectes olivaceus Ochre-bellied Flycatcher Mionectes oleagineus Slaty-capped Flycatcher Leptopogon superciliaris Inca Flycatcher (E) Leptopogon taczanowskii Mottle-cheeked Tyrannulet Phylloscartes ventralis Ecuadorian Tyrannulet Phylloscartes gualaquizae Sooty-headed Tyrannulet Phyllomyias griseiceps Black-capped Tyrannulet Phyllomyias nigrocapillus Red-billed Tyrannulet Zimmerius cinereicapilla Mishana Tyrannulet (E) Zimmerius villarejoi Peruvian Tyrannulet (E) Zimmerius viridiflavus Many-colored Rush Tyrant Tachuris rubrigastra Rufous-headed Pygmy-Tyrant Pseudotriccus ruficeps Tawny-crowned Pygmy-Tyrant Euscarthmus meloryphus Gray-and-white Tyrannulet Pseudelaenia leucospodia H White-bellied Pygmy-Tyrant Myiornis albiventris H Scale-crested Pygmy-Tyrant Lophotriccus pileatus Stripe-necked Tody-Tyrant Hemitriccus striaticollis Pearly-vented Tody-Tyrant Hemitriccus margaritaceiventer Black-throated Tody-Tyrant Hemitriccus granadensis H Cinnamon-breasted Tody-Tyrant Hemitriccus cinnamomeipectus Johnson's Tody-Flycatcher (E) Poecilotriccus luluae Black-and-white Tody-Flycatcher Poecilotriccus capitalis Common Tody-Flycatcher Todirostrum cinereum Yellow-browed Tody-Flycatcher Todirostrum chrysocrotaphum Yellow-margined Flycatcher Tolmomyias assimilis H Yellow-breasted (Olive-faced) Flycatcher Tolmomyias flaviventris White-throated Spadebill Platyrinchus mystaceus Cinnamon Flycatcher Pyrrhomyias cinnamomeus Cliff Flycatcher Hirundinea ferruginea Olive-chested Flycatcher Myiophobus cryptoxanthus